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    building an enduring coexistence of people and threatened ecosystems in sub-Saharan Africa.
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Mountain Nyala (Tragelaphus buxtoni)

The mountain nyala is endemic to Ethiopia (found only in Ethiopia) and is limited in its distribution to the Bale and Arussi provinces of Ethiopia. Because it was the last great African antelope to become to known to science (It was first recorded in 1910), little is known about its habits. It stands 135cms (53 inches) at the shoulder and weighs 200-250 kgs. (440-550 lbs.) and an old bull may weigh as much as 300 kgs. (660 lbs.) As a comparison, elk stand at 122-152 cms (4 to 5 feet) and bull elk weigh 315-450 kgs. (700-1,000 lbs.)

A forest animal, the nyala flourishes in a mosaic of high-altitude woodland (juniper and Hagenia forest), heath, moorland and valley-bottom grassland ranging from approximately 2,100 m to 4,200 m. Plants of the tomato family (Solanaceae), St. John's wort (Hypericum), lady's mantle (Alchemilla) and goosegrass (Galium) are the most frequent browse of the nyala and it is also known to eat fallen leaves of the Hagenia during the dry season. The lower reaches of its range are more suited to the species than the upper less-vegetated areas, but in most parts of its range, this habitat has been taken over for cultivation and pastoralism.

Results of genetic analysis indicate that the mountain nyala is likely to be more closely related to the bushbuck, sitatunga, and the bongo than the Greater Kudu (T. strepsceros) although early accounts referred to the nyala as the Spotted Kudu. However, the male nyala lacks the throat mane and hump on the shoulders of the kudu and is generally smaller in size than its closest relative, weighing up to only 300 kg (660 lbs). Males are a sepia brown color that slowly gets darker with age while females (left) are of a pale liver-color with a scattering of vestigial spots and stripes. Mating peaks in December and single young are born after an 8-9 month gestation, at the end of the wet season. Females accompanied by one or two generations of young form very frequent but essentially impermanent associations with other mother-young groups. A young mountain nyala stays closely attached to its mother for as long as 2 years, by which time female calves are themselves pregnant and males have long horns and join bachelor groups. Older males usually remain solitary.

The Tragelaphini - A Genetic Analysis
(September 2002)

It has been theorized that all African spiral-horned bovines (tragelaphines), which include the genus Tragelaphus and Taurotragus, derive from a single immigrant ancestral type which subsequently branched into a larger and smaller lineage. The relationships of species within the group have been subject to speculation up until now. Genetic analysis of the Tragelaphus species, which include the mountain nyala, bushbuck and greater kudu has recently been conducted by the Smithsonian Institution. This chart, contributed to the newsletter by a member of TMF, shows the results of the analysis.

Contrary to what is generally accepted, that the mountain nyala's (T. buxtoni) closest relatives are the southern nyala and greater kudu, the data suggests that the closest relatives to the mountain nyala are the bushbuck (T. scriptus), sitatunga (T. spekii), and the bongo (T. eurycerus). The results of this analysis are represented in a phylogenetic tree diagram (Fig.1) which uses an index from 0 to 100 (weak to strong) to rank the strength of each association. It should be noted that tissue from the Giant Eland (Tragelaphus spp.) was not available and is not included in the results.

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